This edition fi rst published 2011, © 2011 by Blackwell Publishing Ltd
Blackwell Publishing was acquired by John Wiley & Sons in February 2007. Blackwell’s publishing program has been merged
with Wiley’s global Scientifi c, Technical and Medical business to form Wiley-Blackwell.
Registered offi ce: John Wiley & Sons Ltd, The Atrium, Southern Gate, Chichester, West Sussex, PO19 8SQ, UK
Editorial offi ces: 9600 Garsington Road, Oxford, OX4 2DQ, UK
The Atrium, Southern Gate, Chichester, West Sussex, PO19 8SQ, UK
111 River Street, Hoboken, NJ 07030-5774, USA
For details of our global editorial offi ces, for customer services and for information about how to apply for permission to reuse
the copyright material in this book please see our website at www.wiley.com/wiley-blackwell
The right of the author to be identifi ed as the author of this work has been asserted in accordance with the Copyright, Designs
and Patents Act 1988.
All rights reserved. No part of this publication may be reproduced, stored in a retrieval system, or transmitted, in any form or
by any means, electronic, mechanical, photocopying, recording or otherwise, except as permitted by the UK Copyright, Designs
and Patents Act 1988, without the prior permission of the publisher.
Wiley also publishes its books in a variety of electronic formats. Some content that appears in print may not be available in
electronic books.
Designations used by companies to distinguish their products are often claimed as trademarks. All brand names and product
names used in this book are trade names, service marks, trademarks or registered trademarks of their respective owners. The
publisher is not associated with any product or vendor mentioned in this book. This publication is designed to provide accurate
and authoritative information in regard to the subject matter covered. It is sold on the understanding that the publisher is not
engaged in rendering professional services. If professional advice or other expert assistance is required, the services of a
competent professional should be sought.
Library of Congress Cataloguing-in-Publication Data
Conservation biogeography / edited by R. J. Ladle and R. J. Whittaker.
p. cm.
Includes index.
ISBN 978-1-4443-3503-3 (cloth) – ISBN 978-1-4443-3504-0 (pbk.)
1. Conservation biology. 2. Biodiversity conservation. 3. Protected areas. 4. Biogeography. I. Ladle, Richard J.
II. Whittaker, Robert J.
QH75.C657 2011
333.95′16–dc22
2010037700
A catalogue record for this book is available from the British Library.
This book is published in the following electronic formats: eBook ISBN: 9781444390018; Wiley Online Library ISBN:
9781444390001; ePub ISBN: 9781444390025
Set in 9/11pt Photina by Toppan Best-set Premedia Limited
1 2011
Blackwell Publishing was acquired by John Wiley & Sons in February 2007. Blackwell’s publishing program has been merged
with Wiley’s global Scientifi c, Technical and Medical business to form Wiley-Blackwell.
Registered offi ce: John Wiley & Sons Ltd, The Atrium, Southern Gate, Chichester, West Sussex, PO19 8SQ, UK
Editorial offi ces: 9600 Garsington Road, Oxford, OX4 2DQ, UK
The Atrium, Southern Gate, Chichester, West Sussex, PO19 8SQ, UK
111 River Street, Hoboken, NJ 07030-5774, USA
For details of our global editorial offi ces, for customer services and for information about how to apply for permission to reuse
the copyright material in this book please see our website at www.wiley.com/wiley-blackwell
The right of the author to be identifi ed as the author of this work has been asserted in accordance with the Copyright, Designs
and Patents Act 1988.
All rights reserved. No part of this publication may be reproduced, stored in a retrieval system, or transmitted, in any form or
by any means, electronic, mechanical, photocopying, recording or otherwise, except as permitted by the UK Copyright, Designs
and Patents Act 1988, without the prior permission of the publisher.
Wiley also publishes its books in a variety of electronic formats. Some content that appears in print may not be available in
electronic books.
Designations used by companies to distinguish their products are often claimed as trademarks. All brand names and product
names used in this book are trade names, service marks, trademarks or registered trademarks of their respective owners. The
publisher is not associated with any product or vendor mentioned in this book. This publication is designed to provide accurate
and authoritative information in regard to the subject matter covered. It is sold on the understanding that the publisher is not
engaged in rendering professional services. If professional advice or other expert assistance is required, the services of a
competent professional should be sought.
Library of Congress Cataloguing-in-Publication Data
Conservation biogeography / edited by R. J. Ladle and R. J. Whittaker.
p. cm.
Includes index.
ISBN 978-1-4443-3503-3 (cloth) – ISBN 978-1-4443-3504-0 (pbk.)
1. Conservation biology. 2. Biodiversity conservation. 3. Protected areas. 4. Biogeography. I. Ladle, Richard J.
II. Whittaker, Robert J.
QH75.C657 2011
333.95′16–dc22
2010037700
A catalogue record for this book is available from the British Library.
This book is published in the following electronic formats: eBook ISBN: 9781444390018; Wiley Online Library ISBN:
9781444390001; ePub ISBN: 9781444390025
Set in 9/11pt Photina by Toppan Best-set Premedia Limited
1 2011