pools. Attains 3 cm. East Africa south to Inhaca, Seychelles, Mauritius and St. Brandon's Shoals.
Goggles
240.54 Hetereleotris zonata
(Fowler, 1934)
PLATE 124
Brilletjie
Leioeleotris zonatus Fowler, 1934: 494, Fig. 48 (St. Lucia coast, Natal).
Hetereleotris diadematus (non Riippell): Smith, SFSA No. 935*.
Hetereleotris zonatus: Smith, 1958: 160* 1961: 570, No. 935*.
D VI+ 1,11-13 (11-12 in our area); A 1,9-11 (9-10 in our area); P 16-18;
V 1,5; LS 41-46; TRB 11-13; PD O. H. kenyae Smith, 1958 from east Africa
and Red Sea has 32-36 LS.
Attains 6 cm. Natal to Port Alfred, Pakistan and India.
240.55 Hetereleotris margaretae
Hoese, 1986
Smooth scale goby
Gladdeskub-dikkop
Hetereleotris margaretae Hoese, 1986: 00, Figs. 2 & 3 (Sodwana Bay).
D VI + 1,11; A 1,10; P 17; V 1,5; LS 28-30.
Dark spot above end of opercle; body with about 6 dark
bars; 2 faint dark bars under eye. Attains 3 cm. Known only
from Sodwana Bay.
240.55 Hetereleotris margaretae: 2.5 em SL female holotype (Natal).
Genus Istigobius Whitley, 1932
Mouth small, subinferior; head pores Type A; papillae
pattern longitudinal. Scales ctenoid, none on cheek or operculum. Va disc. Gill opening equals P base. D VI+I,lO;
A 1,9; P 17-20; LS 25-28. Vertebrae 26. Easily confused
with Gnatholepis, which has paired anterior interorbital
pores and cheek scales, with extralimital Exyrias and Macrodontogobius, which have cheek scales and with Oplopomops, which has a thickened stiff spine in 2nd D.
About 10 species, mostly tropical, found in sandy areas
along shore and on coral reefs; 2 species in our area, but a
third, I. ornatus (Riippell, 1830), may be found here.
KEY TO SPECIES
1 a Upper P rays free, silk -like; no enlarged curved
tooth laterally in lower jaw; (mangroves) ..................... l.ornatus
1b No P rays free; an enlarged tooth laterally in outer
row of lower jaw; (reefs) ..................................................... 2
240: GOBIIDAE (GOBlES)
2a Isthmus broad, scaled forward to under eye, scales
extending over branchiostegal membrane;
dusky blotch under eye; no enlarged spot
between 1st 2 D spines .............................................. I. spence
2b Isthmus narrow; scaled forward to behind eye;
cheek with 2 or 3 oblique dark lines; usually with
a black spot between 1st 2 D spines .......................... I. decoratus
240.56 Istigobius decoratus
(Herre, 1927)
Decorated goby
Versierde dikkop
Rhinogobius decoratus Herre, 1927: 181, PI. 13, Fig. 3 (Philippines).
Acentrogobius ornatus (non Riippell): Smith, 1959: 202 (in part).
lstigobius sp.: Hoese & Winterbottom, 1979: 4.
D VI + 1,10; A 1,9; P 17-19; LS 26-27; TRB 7-8; PD 8-10; GR 2+5.
Body yellowish, with dark lines forming a reticulated network dorsally, 4 or 5 pairs of pupil-sized black spots midlaterally, and similar sized spots in a row below; C with enlarged spots; 3 oblique bars below eye. Often confused with
I. ornatus. Attains 13 cm. Indo-West Pacific south to Durban.
240.56 Istigobius decoratus: top: 9 em (Mauritius; photo by J. E. Randall);
bottom: 3.5 em holotype (after Herre, 1927).
240.57 Istiogobius ornatus
(RuppeU, 1830)
Ornate goby
Oorlaide dikkop
Gobius ornatus Riippell, 1830: 135 (Massaua, Red Sea). Acentrogobius
ornatus: Smith, 1959: 202* (in part).
D VI + 1,10; A 1,9; P 17-20; LS 29-32; TRB 8-9; PD 9-12; GR 2+(4-5)
240.57 Istigobius ornatus: 7.5 em (N. Mozambique).
793
Goggles
240.54 Hetereleotris zonata
(Fowler, 1934)
PLATE 124
Brilletjie
Leioeleotris zonatus Fowler, 1934: 494, Fig. 48 (St. Lucia coast, Natal).
Hetereleotris diadematus (non Riippell): Smith, SFSA No. 935*.
Hetereleotris zonatus: Smith, 1958: 160* 1961: 570, No. 935*.
D VI+ 1,11-13 (11-12 in our area); A 1,9-11 (9-10 in our area); P 16-18;
V 1,5; LS 41-46; TRB 11-13; PD O. H. kenyae Smith, 1958 from east Africa
and Red Sea has 32-36 LS.
Attains 6 cm. Natal to Port Alfred, Pakistan and India.
240.55 Hetereleotris margaretae
Hoese, 1986
Smooth scale goby
Gladdeskub-dikkop
Hetereleotris margaretae Hoese, 1986: 00, Figs. 2 & 3 (Sodwana Bay).
D VI + 1,11; A 1,10; P 17; V 1,5; LS 28-30.
Dark spot above end of opercle; body with about 6 dark
bars; 2 faint dark bars under eye. Attains 3 cm. Known only
from Sodwana Bay.
240.55 Hetereleotris margaretae: 2.5 em SL female holotype (Natal).
Genus Istigobius Whitley, 1932
Mouth small, subinferior; head pores Type A; papillae
pattern longitudinal. Scales ctenoid, none on cheek or operculum. Va disc. Gill opening equals P base. D VI+I,lO;
A 1,9; P 17-20; LS 25-28. Vertebrae 26. Easily confused
with Gnatholepis, which has paired anterior interorbital
pores and cheek scales, with extralimital Exyrias and Macrodontogobius, which have cheek scales and with Oplopomops, which has a thickened stiff spine in 2nd D.
About 10 species, mostly tropical, found in sandy areas
along shore and on coral reefs; 2 species in our area, but a
third, I. ornatus (Riippell, 1830), may be found here.
KEY TO SPECIES
1 a Upper P rays free, silk -like; no enlarged curved
tooth laterally in lower jaw; (mangroves) ..................... l.ornatus
1b No P rays free; an enlarged tooth laterally in outer
row of lower jaw; (reefs) ..................................................... 2
240: GOBIIDAE (GOBlES)
2a Isthmus broad, scaled forward to under eye, scales
extending over branchiostegal membrane;
dusky blotch under eye; no enlarged spot
between 1st 2 D spines .............................................. I. spence
2b Isthmus narrow; scaled forward to behind eye;
cheek with 2 or 3 oblique dark lines; usually with
a black spot between 1st 2 D spines .......................... I. decoratus
240.56 Istigobius decoratus
(Herre, 1927)
Decorated goby
Versierde dikkop
Rhinogobius decoratus Herre, 1927: 181, PI. 13, Fig. 3 (Philippines).
Acentrogobius ornatus (non Riippell): Smith, 1959: 202 (in part).
lstigobius sp.: Hoese & Winterbottom, 1979: 4.
D VI + 1,10; A 1,9; P 17-19; LS 26-27; TRB 7-8; PD 8-10; GR 2+5.
Body yellowish, with dark lines forming a reticulated network dorsally, 4 or 5 pairs of pupil-sized black spots midlaterally, and similar sized spots in a row below; C with enlarged spots; 3 oblique bars below eye. Often confused with
I. ornatus. Attains 13 cm. Indo-West Pacific south to Durban.
240.56 Istigobius decoratus: top: 9 em (Mauritius; photo by J. E. Randall);
bottom: 3.5 em holotype (after Herre, 1927).
240.57 Istiogobius ornatus
(RuppeU, 1830)
Ornate goby
Oorlaide dikkop
Gobius ornatus Riippell, 1830: 135 (Massaua, Red Sea). Acentrogobius
ornatus: Smith, 1959: 202* (in part).
D VI + 1,10; A 1,9; P 17-20; LS 29-32; TRB 8-9; PD 9-12; GR 2+(4-5)
240.57 Istigobius ornatus: 7.5 em (N. Mozambique).
793