308
at their basis, widening in their distal parts and almost
joined medially (Fig. 1. E). Ventral lobes on average
smaller than distal ones with conical shape (Fig. 1. F).
Distal article of antenna bent gently inward in its proximal half; proximal end of article with prominent fingerlike outgrowth on medioventral surface (Fig. 1. F). In
area of curving, distal article of antenna bears another several times smaller fingerlike outgrowth on its
ventro-Iateral surface (Fig. 1. F). Tip of article divided in two lobes; dorsal lobe wider and longer than
ventral lobe (Fig. 1. F). Labrum with proximal and
distal lobe. Proximal lobe with conspicuous protuberance on apical part (Fig. 1. C,D). Thoracic appendages
of usual shape and construction as typical of the genus
(Fig. 1. G,H,I). Penis typical of genus (Fig. 1. J). Cercopods long, at least as long as last five postgenital
segments, with plumose setae on lateral and median
borders (Fig. 1. J).
Female Total length, from front to end of shaft of cercopods 9 to 17 mm. Antennula as long as antenna.
Antenna uniarticulate, proximal two thirds cylindrical,
distal third narrowing to tapering chitinized thorn at
apex (Fig. 2. A). Labrum and thoracic appendages as
in male. Penultimate and ultimate (10th and 11th) thoracic segments with one pair of lateral bosses on their
posterior margins (Fig. 2. B,D). Cysts lenticular with
approximate diameter 0.35 to 0.40 mm.
Taxonomic notes
Macedonian specimens are relatively large. Total
length in males 8 to 16.5 mm, in females 9 to 17 mm.
Daday (1910) gave the length of the males as 6.2 to
11.5 mm and the females as 6.2 to 12 mm. Brtek (1962)
gave measurements for the species in general as 6.5 to
12 mm. The specimens examined by Cottarelli & Mura
(1983) measured 9 to 11 mm and Alonso's (1985) specimens from Spain were 6 to 9 mm long. Noteworthy
is the fact that these measurements are much less than
those of the Macedonian specimens. Nevertheless, systematic position of this species is well established and
the Macedonian specimens of T. stagnalis correspond
morphologically to the previous descriptions of the
species from other localities within its range. Furthermore, after this report it seems more probable that
specimens of T. stagnalis can be as large as 20 mm
long, as reported by Nourisson & Thiery (1988) from
France.
Distribution and habitat
According to Brtek (1962) T. stagnalis is referred
to as a Circum-Mediterranean species. The range of
the species stretches from North Africa across Portugal and Spain through France, Switzerland, Austria, Hungary and Bulgaria to Southwestern Russia.
It also extends North across Germany and Denmark
into Scandinavia. T. stagnalis is commonly associated
with small, temporary water bodies. In Macedonia it is
present only on a restricted area in rock pools scattered
over a rocky plain of Quarternary- Pliocene, andesite
tuffs above the village of Stracin in N. E. Macedonia at an elevation of approximately 800 m a.s.l. The
rock pools are extremely small water bodies containing only severallitres of water, measuring 20 to 60 cm
in diameter, and 10 to 20 cm in depth with detritus
of dry leafs, grass, and moss at the bottom. Tanymastix stagnalis is the only anostracan present in these
rock pools, whose advantages include the fact that they
are predator-free and competitor-poor water bodies. In
the neighbouring, larger, temporary water bodies with
soil-substratum T. stagnalis is absent, but there, Chirocephalus diaphanus carinatus, a much larger and competitively more adapted species is numerous.
Tanymastix motasi Orghidan, 1945
Material and locality
EL 1: EL37, Small rain puddles near the village Golemo Konjari, Pelagonia, S. W. Macedonia, 25.05.1985,
31 dd ,47W, leg. S. Petkovski; 26.05.1985,44 dd,
63W, leg. S. Petkovski; 26.04.1991,14 do ,15W,
leg. S. Petkovski; 22.05.1991,129 do ,161 W, leg.
S. Petkovski.
Description
Male On average smaller than female. Total length
from frontto end of shaft of cercopods 7.2 to 14.5 mm.
Antennula conspicuously overreaches distal end of
proximal article of antenna, but in length it never
exceeds whole antenna (Fig. 3. A,B). Antenna biarticulate. Proximal articles of antennae in their basal parts
joined medially, forming clypeus (Fig. 3. A,B,G,H).
Antennal appendages arising from dorsal, proximal
portion of clypeus typical of genus (Fig. 3. A). Mediodistal outgrowth of clypeus with two pairs of chitinized lobes (Fig. 3. G,H). Dorsal lobes widest at their
bases, medially close together, and distally parallel
to each other. Laterally, narrowing in their distal parts
(Fig. 3. G), quite opposite situation in previous species.
at their basis, widening in their distal parts and almost
joined medially (Fig. 1. E). Ventral lobes on average
smaller than distal ones with conical shape (Fig. 1. F).
Distal article of antenna bent gently inward in its proximal half; proximal end of article with prominent fingerlike outgrowth on medioventral surface (Fig. 1. F). In
area of curving, distal article of antenna bears another several times smaller fingerlike outgrowth on its
ventro-Iateral surface (Fig. 1. F). Tip of article divided in two lobes; dorsal lobe wider and longer than
ventral lobe (Fig. 1. F). Labrum with proximal and
distal lobe. Proximal lobe with conspicuous protuberance on apical part (Fig. 1. C,D). Thoracic appendages
of usual shape and construction as typical of the genus
(Fig. 1. G,H,I). Penis typical of genus (Fig. 1. J). Cercopods long, at least as long as last five postgenital
segments, with plumose setae on lateral and median
borders (Fig. 1. J).
Female Total length, from front to end of shaft of cercopods 9 to 17 mm. Antennula as long as antenna.
Antenna uniarticulate, proximal two thirds cylindrical,
distal third narrowing to tapering chitinized thorn at
apex (Fig. 2. A). Labrum and thoracic appendages as
in male. Penultimate and ultimate (10th and 11th) thoracic segments with one pair of lateral bosses on their
posterior margins (Fig. 2. B,D). Cysts lenticular with
approximate diameter 0.35 to 0.40 mm.
Taxonomic notes
Macedonian specimens are relatively large. Total
length in males 8 to 16.5 mm, in females 9 to 17 mm.
Daday (1910) gave the length of the males as 6.2 to
11.5 mm and the females as 6.2 to 12 mm. Brtek (1962)
gave measurements for the species in general as 6.5 to
12 mm. The specimens examined by Cottarelli & Mura
(1983) measured 9 to 11 mm and Alonso's (1985) specimens from Spain were 6 to 9 mm long. Noteworthy
is the fact that these measurements are much less than
those of the Macedonian specimens. Nevertheless, systematic position of this species is well established and
the Macedonian specimens of T. stagnalis correspond
morphologically to the previous descriptions of the
species from other localities within its range. Furthermore, after this report it seems more probable that
specimens of T. stagnalis can be as large as 20 mm
long, as reported by Nourisson & Thiery (1988) from
France.
Distribution and habitat
According to Brtek (1962) T. stagnalis is referred
to as a Circum-Mediterranean species. The range of
the species stretches from North Africa across Portugal and Spain through France, Switzerland, Austria, Hungary and Bulgaria to Southwestern Russia.
It also extends North across Germany and Denmark
into Scandinavia. T. stagnalis is commonly associated
with small, temporary water bodies. In Macedonia it is
present only on a restricted area in rock pools scattered
over a rocky plain of Quarternary- Pliocene, andesite
tuffs above the village of Stracin in N. E. Macedonia at an elevation of approximately 800 m a.s.l. The
rock pools are extremely small water bodies containing only severallitres of water, measuring 20 to 60 cm
in diameter, and 10 to 20 cm in depth with detritus
of dry leafs, grass, and moss at the bottom. Tanymastix stagnalis is the only anostracan present in these
rock pools, whose advantages include the fact that they
are predator-free and competitor-poor water bodies. In
the neighbouring, larger, temporary water bodies with
soil-substratum T. stagnalis is absent, but there, Chirocephalus diaphanus carinatus, a much larger and competitively more adapted species is numerous.
Tanymastix motasi Orghidan, 1945
Material and locality
EL 1: EL37, Small rain puddles near the village Golemo Konjari, Pelagonia, S. W. Macedonia, 25.05.1985,
31 dd ,47W, leg. S. Petkovski; 26.05.1985,44 dd,
63W, leg. S. Petkovski; 26.04.1991,14 do ,15W,
leg. S. Petkovski; 22.05.1991,129 do ,161 W, leg.
S. Petkovski.
Description
Male On average smaller than female. Total length
from frontto end of shaft of cercopods 7.2 to 14.5 mm.
Antennula conspicuously overreaches distal end of
proximal article of antenna, but in length it never
exceeds whole antenna (Fig. 3. A,B). Antenna biarticulate. Proximal articles of antennae in their basal parts
joined medially, forming clypeus (Fig. 3. A,B,G,H).
Antennal appendages arising from dorsal, proximal
portion of clypeus typical of genus (Fig. 3. A). Mediodistal outgrowth of clypeus with two pairs of chitinized lobes (Fig. 3. G,H). Dorsal lobes widest at their
bases, medially close together, and distally parallel
to each other. Laterally, narrowing in their distal parts
(Fig. 3. G), quite opposite situation in previous species.