have yet become involved with reefal bryozoans for such
greater scientific diversification to have developed so far.
Much remains to be done!
Summary
Although initially overlooked by reef scientists, bryozoans living on or in modern coral reefs are common
and diversified on many such structures, particularly in
their principal role as hidden encrusters. Much detailed
work needs to be done yet to fully elaborate many aspects
of reefal bryozoan taxonomy, distribution, and ecology.
Bibliography
Audouin, J.V., 1826. Explication sommaire des planches de polypes
de l’Égypte et de la Syrie, publiées par Jules-César Savigny. In
Description de l’ Égypte, [par] l’Expedition de l’Armée
française; Histoire naturelle. Paris: Imprimérie impériale de sa
Majesté l’Empéreur Napoléon le Grand (republished 1828,
Paris: C.L.F. Panckouche), Vol. 1(4), pp. 225–244.
Bijma, J., and Boekschoten, G. J., 1985. Recent bryozoan reefs and
stromatolite development in brackish inland lakes, SW
Netherlands. Senckenbergiana Maritima, 17, 163–185.
Bone, Y., and Wass, R. E., 1990. Sub–Recent bryozoan-serpulid
buildups in the Coorong Lagoon, South Australia. Australian
Journal of Earth Sciences, 37, 207–214.
Canu, F., and Bassler, R. S., 1928. Fossil and recent bryozoa of the
Gulf of Mexico region. U.S. National Museum Proceedings,
72(14), 1–199.
Canu, F., and Bassler, R. S., 1929. Bryozoa of the Philippine region.
U.S. National Museum Bulletin, 100(9), 1–685.
Cuffey, R. J., 1970. Bryozoan-environment interrelationships – an
overview of bryozoan paleoecology and ecology. Pennsylvania
State University Earth and Mineral Sciences Bulletin, 39(6),
41–45, 48.
Cuffey, R. J., 1972. The roles of bryozoans in modern coral reefs.
Geologische Rundschau, 61, 542–550. (This journal is now
published by Springer as International Journal of Earth
Sciences.)
Cuffey, R. J., 1973. Bryozoan distribution in the modern reefs of
Eniwetok Atoll and the Bermuda Platform. Pacific Geology, 6,
25–50.
Cuffey, R. J., 1977. Bryozoan contributions to reefs and bioherms
through geologic time. American Association of Petroleum
Geologists Studies in Geology, 4, 181–194.
Cuffey, R. J., 1978. A note on bryozoans in the modern reefs of Eniwetok Atoll and the Australian Great Barrier Reef. Pacific Geology, 13, 65–76.
Cuffey, R. J., 1985. Expanded reef-rock textural classification and
the geologic history of bryozoan reefs. Geology, 13, 307–310.
Cuffey, R. J., 2006. Bryozoan-built reef mounds – the overview
from integrating recent studies with previous investigations.
Courier Forschungsinstitut Senckenberg, 257, 35–47.
Cuffey, R. J., and Cox, R. S., 1987. Reef-dwelling bryozoans of
Enewetak Atoll. In Devaney, D. M., Reese, E. S., Burch, B. L.,
and Helfrich, P. (eds.), The Natural History of Enewetak Atoll.
Oak Ridge: U.S. Department of Energy, Vol. 2, pp. 85–92.
Cuffey, R. J., and Foerster, B., 1975. Autecology of the cyclostome
bryozoan Crisia eburnea in the modern reefs of Bermuda.
Bryozoa 1974; Documents des Laboratoires de Géologie de
la Faculté des Sciences de Lyon, hors série 3, Vol. 2, pp.
357–368.
Cuffey, R. J., and Fonda, S. S., 1976. “Giant” Schizoporella – sedimentologically important cheilostome bryozoans in Pleistocene
and Recent carbonate environments of Bermuda, the Bahamas,
and Florida. Geological Society of America Abstracts with Programs, 8, 474–475.
Cuffey, R. J., and Fonda, S. S., 1977. Cryptic bryozoan species
assemblages in modern coral reefs off Andros and
Eleuthera, Bahamas. In Proceedings, International Coral Reef
Symposium, III, Vol. 1, pp. 81–86.
Cuffey, R. J., and Fonda, S. S., 1986. Phylum Bryozoa (sea mats,
moss animals). In Sterrer, W., and Schoepfer-Sterrer, C. (eds.),
Marine Fauna and Flora of Bermuda: A Systematic Guide to
the Identification of Marine Organisms. New York: Wiley, pp.
500–516.
Cuffey, R. J., Gebelein, C. D., Fonda, S. S., Bliefnick, D. M.,
Kosich, D. F., and Soroka, L. G., 1977. Modern tidalchannel bryozoan reefs at Joulters Cays (Bahamas). In
Proceedings, International Coral Reef Symposium, III, Vol. 2,
pp. 339–345.
Cuffey, R. J., and McKinney, F. K., 1982. Reteporid cheilostome
bryozoans from the modern reefs of Enewetak Atoll, and their
implications for Paleozoic fenestrate bryozoan paleoecology.
Pacific Geology, 16, 7–13.
Cuffey, R. J., and Montaggioni, L. F., 1986. Discovery and significance of fossil reefal bryozoans in the uplifted Miocene “atoll” of
Makatea (French Polynesia). Geological Society of America
Abstracts with Programs, 18, 577.
Cuffey, R. J., and Utgaard, J. E., 1999. Bryozoans. In Singer, R.
(ed.), Encyclopedia of Paleontology. Chicago: Fitzroy Dearborn,
Vol. 1, pp. 204–216.
Fonda, S. S., and Cuffey, R. J., 1976. Bryozoan assemblages in the
modern Bermuda reefs – comparative data for analysis of Cenozoic bioherms. Geological Society of America Abstracts with
Programs, 8, 175.
Hamdane, A., and Moissette, P., 2002. Bryozoan distribution
in a Messinian coral reef complex of western Algeria. In Wyse
Jackson, P. N., Buttler, C. J., and Spencer Jones, M. E. (eds.),
Bryozoan Studies 2001. Lisse: A.A. Balkema, pp. 141–149.
Jackson, J. B. C., 1979. Overgrowth competition between
encrusting cheilostome ectoprocts in a Jamaican cryptic reef
environment. Journal of Animal Ecology, 48, 805–823.
Jackson, J. B. C., and Buss, L. W., 1975. Allelopathy and spatial
competition among coral reef invertebrates. Proceedings of the
National Academy of Science of the United States of America,
72, 5160–5163.
Jackson, J. B. C., and Winston, J. E., 1982. Ecology of coral reef
communities. I. Distribution and abundance of major groups.
Journal of Experimental Marine Biology, 57, 135–147.
Kobluk, D. R., Cuffey, R. J., Fonda, S. S., and Lysenko, M. A.,
1988. Cryptic Bryozoa, leeward fringing reef of Bonaire,
Netherlands Antilles, and their paleoecological application.
Journal of Paleontology, 62, 427–439.
Kosich, D. F., and Cuffey, R. J., 1978. Recognition of multilaminar
cheilostome bryozoan species in modern Bermuda reefrock. Geological Society of America Abstracts with Programs,
10, 259.
Maxwell, W. G. H., 1968. Atlas of the Great Barrier Reef.
Amsterdam: Elsevier.
Ortmann, A., 1892. Die Koralriffe von Dar-es-Salaam und
Umgegend. Zoologische Jahrbuch, Abteilung fűr Systematik,
6, 631–670.
Osburn, R. C., 1940. Bryozoa of Porto Rico with a résumé of the
West Indian bryozoan fauna. New York Academy of Sciences Scientific Survey of Porto Rico and the Virgin Islands, 16(3),
321–486.
Pitcher, M., 1964. Evolution of Chazyan (Ordovician) reefs of eastern United States and Canada. Bulletin of Canadian Petroleum
Geology, 12, 632–691.
176
BRYOZOA
greater scientific diversification to have developed so far.
Much remains to be done!
Summary
Although initially overlooked by reef scientists, bryozoans living on or in modern coral reefs are common
and diversified on many such structures, particularly in
their principal role as hidden encrusters. Much detailed
work needs to be done yet to fully elaborate many aspects
of reefal bryozoan taxonomy, distribution, and ecology.
Bibliography
Audouin, J.V., 1826. Explication sommaire des planches de polypes
de l’Égypte et de la Syrie, publiées par Jules-César Savigny. In
Description de l’ Égypte, [par] l’Expedition de l’Armée
française; Histoire naturelle. Paris: Imprimérie impériale de sa
Majesté l’Empéreur Napoléon le Grand (republished 1828,
Paris: C.L.F. Panckouche), Vol. 1(4), pp. 225–244.
Bijma, J., and Boekschoten, G. J., 1985. Recent bryozoan reefs and
stromatolite development in brackish inland lakes, SW
Netherlands. Senckenbergiana Maritima, 17, 163–185.
Bone, Y., and Wass, R. E., 1990. Sub–Recent bryozoan-serpulid
buildups in the Coorong Lagoon, South Australia. Australian
Journal of Earth Sciences, 37, 207–214.
Canu, F., and Bassler, R. S., 1928. Fossil and recent bryozoa of the
Gulf of Mexico region. U.S. National Museum Proceedings,
72(14), 1–199.
Canu, F., and Bassler, R. S., 1929. Bryozoa of the Philippine region.
U.S. National Museum Bulletin, 100(9), 1–685.
Cuffey, R. J., 1970. Bryozoan-environment interrelationships – an
overview of bryozoan paleoecology and ecology. Pennsylvania
State University Earth and Mineral Sciences Bulletin, 39(6),
41–45, 48.
Cuffey, R. J., 1972. The roles of bryozoans in modern coral reefs.
Geologische Rundschau, 61, 542–550. (This journal is now
published by Springer as International Journal of Earth
Sciences.)
Cuffey, R. J., 1973. Bryozoan distribution in the modern reefs of
Eniwetok Atoll and the Bermuda Platform. Pacific Geology, 6,
25–50.
Cuffey, R. J., 1977. Bryozoan contributions to reefs and bioherms
through geologic time. American Association of Petroleum
Geologists Studies in Geology, 4, 181–194.
Cuffey, R. J., 1978. A note on bryozoans in the modern reefs of Eniwetok Atoll and the Australian Great Barrier Reef. Pacific Geology, 13, 65–76.
Cuffey, R. J., 1985. Expanded reef-rock textural classification and
the geologic history of bryozoan reefs. Geology, 13, 307–310.
Cuffey, R. J., 2006. Bryozoan-built reef mounds – the overview
from integrating recent studies with previous investigations.
Courier Forschungsinstitut Senckenberg, 257, 35–47.
Cuffey, R. J., and Cox, R. S., 1987. Reef-dwelling bryozoans of
Enewetak Atoll. In Devaney, D. M., Reese, E. S., Burch, B. L.,
and Helfrich, P. (eds.), The Natural History of Enewetak Atoll.
Oak Ridge: U.S. Department of Energy, Vol. 2, pp. 85–92.
Cuffey, R. J., and Foerster, B., 1975. Autecology of the cyclostome
bryozoan Crisia eburnea in the modern reefs of Bermuda.
Bryozoa 1974; Documents des Laboratoires de Géologie de
la Faculté des Sciences de Lyon, hors série 3, Vol. 2, pp.
357–368.
Cuffey, R. J., and Fonda, S. S., 1976. “Giant” Schizoporella – sedimentologically important cheilostome bryozoans in Pleistocene
and Recent carbonate environments of Bermuda, the Bahamas,
and Florida. Geological Society of America Abstracts with Programs, 8, 474–475.
Cuffey, R. J., and Fonda, S. S., 1977. Cryptic bryozoan species
assemblages in modern coral reefs off Andros and
Eleuthera, Bahamas. In Proceedings, International Coral Reef
Symposium, III, Vol. 1, pp. 81–86.
Cuffey, R. J., and Fonda, S. S., 1986. Phylum Bryozoa (sea mats,
moss animals). In Sterrer, W., and Schoepfer-Sterrer, C. (eds.),
Marine Fauna and Flora of Bermuda: A Systematic Guide to
the Identification of Marine Organisms. New York: Wiley, pp.
500–516.
Cuffey, R. J., Gebelein, C. D., Fonda, S. S., Bliefnick, D. M.,
Kosich, D. F., and Soroka, L. G., 1977. Modern tidalchannel bryozoan reefs at Joulters Cays (Bahamas). In
Proceedings, International Coral Reef Symposium, III, Vol. 2,
pp. 339–345.
Cuffey, R. J., and McKinney, F. K., 1982. Reteporid cheilostome
bryozoans from the modern reefs of Enewetak Atoll, and their
implications for Paleozoic fenestrate bryozoan paleoecology.
Pacific Geology, 16, 7–13.
Cuffey, R. J., and Montaggioni, L. F., 1986. Discovery and significance of fossil reefal bryozoans in the uplifted Miocene “atoll” of
Makatea (French Polynesia). Geological Society of America
Abstracts with Programs, 18, 577.
Cuffey, R. J., and Utgaard, J. E., 1999. Bryozoans. In Singer, R.
(ed.), Encyclopedia of Paleontology. Chicago: Fitzroy Dearborn,
Vol. 1, pp. 204–216.
Fonda, S. S., and Cuffey, R. J., 1976. Bryozoan assemblages in the
modern Bermuda reefs – comparative data for analysis of Cenozoic bioherms. Geological Society of America Abstracts with
Programs, 8, 175.
Hamdane, A., and Moissette, P., 2002. Bryozoan distribution
in a Messinian coral reef complex of western Algeria. In Wyse
Jackson, P. N., Buttler, C. J., and Spencer Jones, M. E. (eds.),
Bryozoan Studies 2001. Lisse: A.A. Balkema, pp. 141–149.
Jackson, J. B. C., 1979. Overgrowth competition between
encrusting cheilostome ectoprocts in a Jamaican cryptic reef
environment. Journal of Animal Ecology, 48, 805–823.
Jackson, J. B. C., and Buss, L. W., 1975. Allelopathy and spatial
competition among coral reef invertebrates. Proceedings of the
National Academy of Science of the United States of America,
72, 5160–5163.
Jackson, J. B. C., and Winston, J. E., 1982. Ecology of coral reef
communities. I. Distribution and abundance of major groups.
Journal of Experimental Marine Biology, 57, 135–147.
Kobluk, D. R., Cuffey, R. J., Fonda, S. S., and Lysenko, M. A.,
1988. Cryptic Bryozoa, leeward fringing reef of Bonaire,
Netherlands Antilles, and their paleoecological application.
Journal of Paleontology, 62, 427–439.
Kosich, D. F., and Cuffey, R. J., 1978. Recognition of multilaminar
cheilostome bryozoan species in modern Bermuda reefrock. Geological Society of America Abstracts with Programs,
10, 259.
Maxwell, W. G. H., 1968. Atlas of the Great Barrier Reef.
Amsterdam: Elsevier.
Ortmann, A., 1892. Die Koralriffe von Dar-es-Salaam und
Umgegend. Zoologische Jahrbuch, Abteilung fűr Systematik,
6, 631–670.
Osburn, R. C., 1940. Bryozoa of Porto Rico with a résumé of the
West Indian bryozoan fauna. New York Academy of Sciences Scientific Survey of Porto Rico and the Virgin Islands, 16(3),
321–486.
Pitcher, M., 1964. Evolution of Chazyan (Ordovician) reefs of eastern United States and Canada. Bulletin of Canadian Petroleum
Geology, 12, 632–691.
176
BRYOZOA