VI II
Contents
8
Grid-Based Modelling of Macrozoobenthos in the Intertidal of the
Wadden Sea: Potentials and Limitations ............................................... 207
V. Grimm, c.-P. Gunther, S. Dittmann & H. Hildenbrandt
8.1
Introduction .................................................................................... 208
8.1.1 Theoretical Background ...................................................... 209
8.1.2 Empirical Background ......................................................... 209
8.2
The First Model: A Demonstration ................................................. 210
8.2.1 Results of the First Model ................................................... 212
8.3
Problems When Applying the Grid-Based Approach in the
Wadden Sea .................................................................................... 214
8.4
A Grid-Based Approach Tailored to the Wadden Sea:
TOPOGRID ....................................................................................... 217
8.4.1 Typical Results of TOPOGRID ................................................ 221
8.5
Methodological Conclusions .......................................................... 223
8.6
Ecological Conclusions ................................................................... 224
9
Stability Properties in the Wadden Sea .................................................. 227
V. Grimm, H. Bietz, c.-P. Gunther, A. Hild, M. Villbrandt, V. Niesel,
U. Schleier & S. Dittmann
9.1
Introduction .................................................................................... 227
9.2
The Conceptual Framework: A Glossary ........................................ 228
9.3
A Canon of Questions ..................................................................... 232
9.4
The Abiotic Ingredients of the Wadden Sea ................................... 234
9.4.1 Meteorological Boundary Conditions and Disturbance
Regime ................................................................................ 234
9.4.2 Hydrography, Morphology and Sediments .......................... 236
9.4.3 Chemistry of the Sediments ................................................. 240
9.4.4 Summary: Abiotic Ingredients of the Wadden Sea .............. 242
9.5 The Biotic Ingredients of the Wadden Sea ..................................... 243
9.5.1 Bacteria ............................................................................... 243
9.5.2 Microphytobenthos and Phytoplankton ............................... 244
9.5.3 Macrozoobenthos ................................................................ 246
9.5.4 Migratory Birds ................................................................... 252
9.5.5 Summary: Biotic Ingredients of the Wadden Sea ................ 253
9.6
Conclusions .................................................................................... 254
10 Assessing Stability Properties: How Suitable is this Approach for
Ecosystem Research in the Wadden Sea? .............................................. 261
S. Dittmann
11 Joint Research Projects: Experiences and Recommendations ............. 267
S. Dittmann, A. Hild, V. Grimm, c.-P. Gunther, V. Niesel,
M. Villbrandt, H. Bietz & U. Schleier
11.1 Realization of ELA W AT ................................................................ 267
11.2 Problems and Potentials of Joint Projects ....................................... 273
Contents
8
Grid-Based Modelling of Macrozoobenthos in the Intertidal of the
Wadden Sea: Potentials and Limitations ............................................... 207
V. Grimm, c.-P. Gunther, S. Dittmann & H. Hildenbrandt
8.1
Introduction .................................................................................... 208
8.1.1 Theoretical Background ...................................................... 209
8.1.2 Empirical Background ......................................................... 209
8.2
The First Model: A Demonstration ................................................. 210
8.2.1 Results of the First Model ................................................... 212
8.3
Problems When Applying the Grid-Based Approach in the
Wadden Sea .................................................................................... 214
8.4
A Grid-Based Approach Tailored to the Wadden Sea:
TOPOGRID ....................................................................................... 217
8.4.1 Typical Results of TOPOGRID ................................................ 221
8.5
Methodological Conclusions .......................................................... 223
8.6
Ecological Conclusions ................................................................... 224
9
Stability Properties in the Wadden Sea .................................................. 227
V. Grimm, H. Bietz, c.-P. Gunther, A. Hild, M. Villbrandt, V. Niesel,
U. Schleier & S. Dittmann
9.1
Introduction .................................................................................... 227
9.2
The Conceptual Framework: A Glossary ........................................ 228
9.3
A Canon of Questions ..................................................................... 232
9.4
The Abiotic Ingredients of the Wadden Sea ................................... 234
9.4.1 Meteorological Boundary Conditions and Disturbance
Regime ................................................................................ 234
9.4.2 Hydrography, Morphology and Sediments .......................... 236
9.4.3 Chemistry of the Sediments ................................................. 240
9.4.4 Summary: Abiotic Ingredients of the Wadden Sea .............. 242
9.5 The Biotic Ingredients of the Wadden Sea ..................................... 243
9.5.1 Bacteria ............................................................................... 243
9.5.2 Microphytobenthos and Phytoplankton ............................... 244
9.5.3 Macrozoobenthos ................................................................ 246
9.5.4 Migratory Birds ................................................................... 252
9.5.5 Summary: Biotic Ingredients of the Wadden Sea ................ 253
9.6
Conclusions .................................................................................... 254
10 Assessing Stability Properties: How Suitable is this Approach for
Ecosystem Research in the Wadden Sea? .............................................. 261
S. Dittmann
11 Joint Research Projects: Experiences and Recommendations ............. 267
S. Dittmann, A. Hild, V. Grimm, c.-P. Gunther, V. Niesel,
M. Villbrandt, H. Bietz & U. Schleier
11.1 Realization of ELA W AT ................................................................ 267
11.2 Problems and Potentials of Joint Projects ....................................... 273