VI
Contents
4.3
Statistical Methods for the Description and Measurement
of Stability Properties ....................................................................... 55
4.3.1 Description of Patterns and Processes ................................... 55
4.3.2 Analysis of Relations ............................................................. 69
4.3.3 Tests for Statistical Distinction Between States of
a System ............................................................................... 70
4.4
Statistics in ELA WAT ...................................................................... 73
5
Spatial and Temporal Distribution Patterns and Their
Underlying Causes ..................................................................................... 77
5.1
Distribution of Nutrients, Algae and Zooplankton in the
Spiekeroog Backbarrier System ......................................................... 77
V. Niesel C. -Po Gunther
5.1.1 Introduction ........................................................................... 77
5.1.2 Material and Methods ............................................................ 78
5.1.3 Tidal Variations ..................................................................... 79
5.1.4 Seasonal Variability .............................................................. 82
5.1.5 Interannual Variability .......................................................... 86
5.1.6 Relations Between Phytoplankton and Copepods .................. 87
5.1.7 Relations Between Phytoplankton and Meroplanktonic Larvae .................................................................. 88
5.1.8 Resuspension of Benthic Algae ............................................. 91
5.2
Biogeochemical Processes in Tidal Flat Sediments and Mutual
Interactions with Macrobenthos ......................................................... 95
M. Villbrandt, A. Hild & S. Dittmann
5.2.1 Microbial and Geochemical Processes .................................. 96
5.2.2 Distribution of Macrofauna Along the Transect .................. 118
5.2.3 Discussion ........................................................................... 123
5.2.4 Conclusions ......................................................................... 128
5.3
Settlement, Secondary Dispersal and Turnover Rate of Benthic
Macrofauna ..................................................................................... 133
C. -P. Gunther
5.3.1 Introduction ......................................................................... 133
5.3.2 What Determines Year-Class Strength in Intertidal Soft
Bottom Benthic Macrofauna? .............................................. 134
5.3.3 Development of Spatial And Temporal Patterns by Initial
Settlement and Secondary Dispersal ................................... 135
5.3.4 In situ Turnover Rates of Juvenile Macrofauna in Soft
Sediments ............................................................................ 141
5.3.5 Small-Scale Temporal Variability in Benthos Distribution
Patterns: Patchiness vS. Mobility .......................................... 142
5.3.6 Lanice conchilega ................................................................ 143
5.4
Modelling the Spatial and Temporal Distribution of Lanice
conchilega ....................................................................................... 147
V. Grimm
Contents
4.3
Statistical Methods for the Description and Measurement
of Stability Properties ....................................................................... 55
4.3.1 Description of Patterns and Processes ................................... 55
4.3.2 Analysis of Relations ............................................................. 69
4.3.3 Tests for Statistical Distinction Between States of
a System ............................................................................... 70
4.4
Statistics in ELA WAT ...................................................................... 73
5
Spatial and Temporal Distribution Patterns and Their
Underlying Causes ..................................................................................... 77
5.1
Distribution of Nutrients, Algae and Zooplankton in the
Spiekeroog Backbarrier System ......................................................... 77
V. Niesel C. -Po Gunther
5.1.1 Introduction ........................................................................... 77
5.1.2 Material and Methods ............................................................ 78
5.1.3 Tidal Variations ..................................................................... 79
5.1.4 Seasonal Variability .............................................................. 82
5.1.5 Interannual Variability .......................................................... 86
5.1.6 Relations Between Phytoplankton and Copepods .................. 87
5.1.7 Relations Between Phytoplankton and Meroplanktonic Larvae .................................................................. 88
5.1.8 Resuspension of Benthic Algae ............................................. 91
5.2
Biogeochemical Processes in Tidal Flat Sediments and Mutual
Interactions with Macrobenthos ......................................................... 95
M. Villbrandt, A. Hild & S. Dittmann
5.2.1 Microbial and Geochemical Processes .................................. 96
5.2.2 Distribution of Macrofauna Along the Transect .................. 118
5.2.3 Discussion ........................................................................... 123
5.2.4 Conclusions ......................................................................... 128
5.3
Settlement, Secondary Dispersal and Turnover Rate of Benthic
Macrofauna ..................................................................................... 133
C. -P. Gunther
5.3.1 Introduction ......................................................................... 133
5.3.2 What Determines Year-Class Strength in Intertidal Soft
Bottom Benthic Macrofauna? .............................................. 134
5.3.3 Development of Spatial And Temporal Patterns by Initial
Settlement and Secondary Dispersal ................................... 135
5.3.4 In situ Turnover Rates of Juvenile Macrofauna in Soft
Sediments ............................................................................ 141
5.3.5 Small-Scale Temporal Variability in Benthos Distribution
Patterns: Patchiness vS. Mobility .......................................... 142
5.3.6 Lanice conchilega ................................................................ 143
5.4
Modelling the Spatial and Temporal Distribution of Lanice
conchilega ....................................................................................... 147
V. Grimm