5.6 Size Frequency. Distribution and Colour Variation of C{{(('inlls l1Iael1llS
173
McGaw IJ. Kaiser MJ, Naylor E. Hughes RN (1992) Intraspccific morphological variation
related to the moult-cycle in color forms of the shore crab Carcinus l1Iaenas. J Zool 228:
351-359
McGaw IJ. Naylor E (19Y2a) Distribution and rhythmic locomotor patterns of estuarine and
open-shore populations of Carcinus maenas. J Mar Bioi Ass UK 72: 59Y-609
McGaw IJ, Naylor E (1992b) The effect of shelter on salinity preference behaviour of the shore
crab Carcinu.I' maena.l'. Mar Behav Physiol 21: 145-152
Moore, PG & J Howarth (I Y(6) Foraging by marine scavengers: effects of relatedness, bait
damage and hunger. 1. Sea Res 36: 267-273.
Moksnes PO, Pihl L, van Montfrans J (I YY8) Predation on postlarvae and juveniles of the shore
crab CarcinLis maenas: importance of shelter, size and cannibalism. Mar Ecol Prog Ser 166:
211-225
Munch-Petersen S, Spane p, Hoffmann E (1982) Abundance of the shore crab, CarclnLl.l' maenas (L.), estimated from mark-recapture experiments. Dana 2: 97-121
Naylor E (1962) Seasonal changes in a population of CarClnus maenas in the littoral zone. J
Animal Ecol 31: 601-609
Rangeley RW, Thomas MLH (1987) Predatory behaviour of the juvenile shore crab CarClntlS
maenas. J Exp Mar Bioi Ecol 108: 191-197
Reid DG. Aldrich JC (1989) Variations in response to environmental hypoxia of different color
forms of the shore crab. C([rcinus maenas. Comp Biochem Pysiol 92: 535-539
Reid DG, Abello p, McGaw /J, Naylor E (1989) Phenotypic variation in sympatric crab populations. In: Aldrich JC (Ed) Phenotypic responses and individuality in aquatic ectoterms. Co.
Wicklow, Ireland, Japaga. pp 8Y-95
Reid DG, Abello p, Kaiser MJ. Warman CG (1997) Carapace colour, inter-moult duration and
the behavioural and physiological ecology of the shore crab CarcinLis maenas. Est Coast
Shelf Sci 44: 203-21 I
Reise K (1 (85) Tidal flat ecology. Springer, Berlin Heidelberg New York
Ropes JW (1968) The feeding habits of the green crab Carcinus maenas. Fish Bull Fish Wildl
Serv US 67: 183-203
Scherer B, Reise K (I (81) Significant predation on micro- and macrobenthos of the Wadden
Sea. Kieler Meeresforsch. Sonderheft 5: 490-500
Thiel M, Dernedde T (I YY4) Recruitment of shore crabs (Carclnlls maenas) on tidal nats: mussel clumps as an important refuge forjuveniles. Helgolander Meeresunters 48: 321-332
Walne PR, Dean (I (72) Experiments on predation by the shore crab, Carcinus maenas, on
Mytllll.1 and Mercenaria. J Cons Perm Int Expl Mer 34: 190-J9Y
Warman CG, Reid DG, Naylor E (IYY3) Variations in the tidal migratory behaviour and rhythmic light-responsiveness in the shore crab, Carcintls maen(lS. J Mar Bioi Ass UK 73: 355364
Wolf F (1997) Untersuchungen zum Auftreten roter und gri.iner Farbvarianten bei der
Strandkrabbe, CarcinllS muenus (Linnaeus 1758) (Crustacea: Decapoda: Brachyura). Dissertation, Universitat Frankfurt a. M.
Wolf F (I (98) Red and green colour forms in the common shore crab Carclnlls maenas (L.)
(Crustacea: Brachyura: Portunidae): theoretical predictions and empirical data. Journal of
Natural History 32: 1807-1812
van der Meeren Gro I. (1994) Sex- and size-dependent mating tactics in a natural population of
shore crabs Carcinus !lweI/as. Journal of Animal Ecology 63: 307-314,
173
McGaw IJ. Kaiser MJ, Naylor E. Hughes RN (1992) Intraspccific morphological variation
related to the moult-cycle in color forms of the shore crab Carcinus l1Iaenas. J Zool 228:
351-359
McGaw IJ. Naylor E (19Y2a) Distribution and rhythmic locomotor patterns of estuarine and
open-shore populations of Carcinus maenas. J Mar Bioi Ass UK 72: 59Y-609
McGaw IJ, Naylor E (1992b) The effect of shelter on salinity preference behaviour of the shore
crab Carcinu.I' maena.l'. Mar Behav Physiol 21: 145-152
Moore, PG & J Howarth (I Y(6) Foraging by marine scavengers: effects of relatedness, bait
damage and hunger. 1. Sea Res 36: 267-273.
Moksnes PO, Pihl L, van Montfrans J (I YY8) Predation on postlarvae and juveniles of the shore
crab CarcinLis maenas: importance of shelter, size and cannibalism. Mar Ecol Prog Ser 166:
211-225
Munch-Petersen S, Spane p, Hoffmann E (1982) Abundance of the shore crab, CarclnLl.l' maenas (L.), estimated from mark-recapture experiments. Dana 2: 97-121
Naylor E (1962) Seasonal changes in a population of CarClnus maenas in the littoral zone. J
Animal Ecol 31: 601-609
Rangeley RW, Thomas MLH (1987) Predatory behaviour of the juvenile shore crab CarClntlS
maenas. J Exp Mar Bioi Ecol 108: 191-197
Reid DG. Aldrich JC (1989) Variations in response to environmental hypoxia of different color
forms of the shore crab. C([rcinus maenas. Comp Biochem Pysiol 92: 535-539
Reid DG, Abello p, McGaw /J, Naylor E (1989) Phenotypic variation in sympatric crab populations. In: Aldrich JC (Ed) Phenotypic responses and individuality in aquatic ectoterms. Co.
Wicklow, Ireland, Japaga. pp 8Y-95
Reid DG, Abello p, Kaiser MJ. Warman CG (1997) Carapace colour, inter-moult duration and
the behavioural and physiological ecology of the shore crab CarcinLis maenas. Est Coast
Shelf Sci 44: 203-21 I
Reise K (1 (85) Tidal flat ecology. Springer, Berlin Heidelberg New York
Ropes JW (1968) The feeding habits of the green crab Carcinus maenas. Fish Bull Fish Wildl
Serv US 67: 183-203
Scherer B, Reise K (I (81) Significant predation on micro- and macrobenthos of the Wadden
Sea. Kieler Meeresforsch. Sonderheft 5: 490-500
Thiel M, Dernedde T (I YY4) Recruitment of shore crabs (Carclnlls maenas) on tidal nats: mussel clumps as an important refuge forjuveniles. Helgolander Meeresunters 48: 321-332
Walne PR, Dean (I (72) Experiments on predation by the shore crab, Carcinus maenas, on
Mytllll.1 and Mercenaria. J Cons Perm Int Expl Mer 34: 190-J9Y
Warman CG, Reid DG, Naylor E (IYY3) Variations in the tidal migratory behaviour and rhythmic light-responsiveness in the shore crab, Carcintls maen(lS. J Mar Bioi Ass UK 73: 355364
Wolf F (1997) Untersuchungen zum Auftreten roter und gri.iner Farbvarianten bei der
Strandkrabbe, CarcinllS muenus (Linnaeus 1758) (Crustacea: Decapoda: Brachyura). Dissertation, Universitat Frankfurt a. M.
Wolf F (I (98) Red and green colour forms in the common shore crab Carclnlls maenas (L.)
(Crustacea: Brachyura: Portunidae): theoretical predictions and empirical data. Journal of
Natural History 32: 1807-1812
van der Meeren Gro I. (1994) Sex- and size-dependent mating tactics in a natural population of
shore crabs Carcinus !lweI/as. Journal of Animal Ecology 63: 307-314,