Population Dynamics of Benthic Species and Shorebird Predation
335
Wiersma P, Pier sma T (1994) Effects of microhabitat, flocking, climate and migratory
goal on energy expenditure in the annual cycle of knots. Condor 96:257-279
Wilson WHJ (1991) The foraging ecology of migratory shorebirds in marine soft-sediment communities: the effects of episodic predation on prey populations. Am Zool
31:840-848
Wolff WJ (1991) The interaction of benthic macrofauna and birds in tidal flat estuaries:
a comparison of Banc d' Arguin, Mauritania, and some estuaries in the Netherlands.
In: Elliot M, Ducrotoy JP (eds) Proceedings 19th ECSA Symp Int Symp Ser
Wolff WJ, De Wolf L (1977) Biomass and production of zoobenthos in the Grevelingen
Estuary, The Netherlands. Estuarine Coastal Mar Sci 5:1-24
WolffWJ, Smit CJ (1990) The Banc d'Arguin, Mauritania, as an environment for coastal
birds. Ardea 78:17-38
Wolff WJ, Van Haperen AMM, Sandee AJJ, Baptist HJM, Saeijs HLF (1976) The trophic
role of birds in the Grevelingen estuary, The Netherlands, as compared to their role in
the saline Lake Grevelingen. In: Persoone G, Jaspers E (eds) Proceedings of the 10th
European Symposium on Marine Biology. Universa Press, Wetteren
WolffWJ, Duiven AG, Duiven P, Esselink P, Gueye A, Meijboom A, Moerland G, Zegers J
(1993) Biomass of macrobenthic tidal flat fauna of the Banc d'Arguin, Mauritania.
Hydrobiologia 258:151-163
Yodzis P (1988) The indeterminacy of ecological interactions as perceived through
perturbation experiments. Ecology 69:508-515
Zwarts L (1991) Seasonal variation in body weight of the bivalves Macoma balthica,
Scrobicularia plana, Mya arena ria and Cerastoderma edule in the Dutch Wadden Sea.
Neth J Sea Res 28:231-245
Zwarts L, Blomert AM (1992) Why knots Calidris canutus take medium sized Macoma
balthica when six prey species are available. Mar Ecol Prog Ser 83:113-128
Zwarts L, Drent RH (1981) Prey depletion and the regulation of predator density: oystercatchers (Haematopus ostralegus) feeding on mussels (Mytilus edulis). In: Jones NV,
Wolff WJ (eds) Feeding and survival strategies of estuarine organisms. Plenum,
London
Zwarts L, Blomert AM, Wan ink JH (1992) Annual and seasonal variation in the food
supply harvestable by knots Calidris canutus staging in the Wadden Sea in late
summer. Mar Ecol Prog Ser 83:129-139
Zwarts L, Ens BJ, Goss-Custard JD, Hulscher JB, Kersten M (1996) Why oystercatchers
Haematopus ostralegus cannot meet their daily energy requirements in a single low
water period. Ardea 84A:269-290
335
Wiersma P, Pier sma T (1994) Effects of microhabitat, flocking, climate and migratory
goal on energy expenditure in the annual cycle of knots. Condor 96:257-279
Wilson WHJ (1991) The foraging ecology of migratory shorebirds in marine soft-sediment communities: the effects of episodic predation on prey populations. Am Zool
31:840-848
Wolff WJ (1991) The interaction of benthic macrofauna and birds in tidal flat estuaries:
a comparison of Banc d' Arguin, Mauritania, and some estuaries in the Netherlands.
In: Elliot M, Ducrotoy JP (eds) Proceedings 19th ECSA Symp Int Symp Ser
Wolff WJ, De Wolf L (1977) Biomass and production of zoobenthos in the Grevelingen
Estuary, The Netherlands. Estuarine Coastal Mar Sci 5:1-24
WolffWJ, Smit CJ (1990) The Banc d'Arguin, Mauritania, as an environment for coastal
birds. Ardea 78:17-38
Wolff WJ, Van Haperen AMM, Sandee AJJ, Baptist HJM, Saeijs HLF (1976) The trophic
role of birds in the Grevelingen estuary, The Netherlands, as compared to their role in
the saline Lake Grevelingen. In: Persoone G, Jaspers E (eds) Proceedings of the 10th
European Symposium on Marine Biology. Universa Press, Wetteren
WolffWJ, Duiven AG, Duiven P, Esselink P, Gueye A, Meijboom A, Moerland G, Zegers J
(1993) Biomass of macrobenthic tidal flat fauna of the Banc d'Arguin, Mauritania.
Hydrobiologia 258:151-163
Yodzis P (1988) The indeterminacy of ecological interactions as perceived through
perturbation experiments. Ecology 69:508-515
Zwarts L (1991) Seasonal variation in body weight of the bivalves Macoma balthica,
Scrobicularia plana, Mya arena ria and Cerastoderma edule in the Dutch Wadden Sea.
Neth J Sea Res 28:231-245
Zwarts L, Blomert AM (1992) Why knots Calidris canutus take medium sized Macoma
balthica when six prey species are available. Mar Ecol Prog Ser 83:113-128
Zwarts L, Drent RH (1981) Prey depletion and the regulation of predator density: oystercatchers (Haematopus ostralegus) feeding on mussels (Mytilus edulis). In: Jones NV,
Wolff WJ (eds) Feeding and survival strategies of estuarine organisms. Plenum,
London
Zwarts L, Blomert AM, Wan ink JH (1992) Annual and seasonal variation in the food
supply harvestable by knots Calidris canutus staging in the Wadden Sea in late
summer. Mar Ecol Prog Ser 83:129-139
Zwarts L, Ens BJ, Goss-Custard JD, Hulscher JB, Kersten M (1996) Why oystercatchers
Haematopus ostralegus cannot meet their daily energy requirements in a single low
water period. Ardea 84A:269-290