Andrea Megela Simmons
Arthur N. Popper
Departments of Psychology and
Department of Biology and
Neuroscience
Neuroscience and
Brown University
Cognitive Science Program
Box 1853
University of Maryland
Providence RI 02912, USA
College Park, MD 20742-4415, USA
Andrea_Simmons@brown.edu
AP17@umail.umd.edu
Richard R. Fay
Department of Psychology and
Parmly Hearing Institute
Loyola University of Chicago
Chicago, IL 60626, USA
rfay@wpo.it.luc.edu
Series Editors: Richard R. Fay and Arthur N. Popper
Cover illustration: Courtesy of Daniel Baleckaitis and Daniel Margoliash.
Library of Congress Cataloging-in-Publication Data
Acoustic communication / editors, Andrea Megela Simmons, Arthur N. Popper, Richard
R. Fay.
p. cm.—(Springer handbook of auditory research; 16)
Includes bibliographical references.
ISBN 0-387-98661-8 (alk. paper)
1. Animal sounds. 2. Sound production by animals. 3. Animal communication.
I. Simmons, Andrea Megela. II. Popper, Arthur N. III. Fay, Richard R. IV. Springer
handbook of auditory research; v. 16.
QL765 .A32 2002
591.59¢4—dc21
2001057680
ISBN 0-387-98661-8
Printed on acid-free paper.
© 2003 Springer-Verlag New York, Inc.
All rights reserved. This work may not be translated or copied in whole or in part without the
written permission of the publisher (Springer-Verlag New York, Inc., 175 Fifth Avenue, New
York, NY 10010, USA), except for brief excerpts in connection with reviews or scholarly
analysis. Use in connection with any form of information storage and retrieval, electronic
adaptation, computer software, or by similar or dissimilar methodology now know or hereafter developed is forbidden.
The use in this publication of trade names, trademarks, service marks, and similar terms, even
if they are not identified as such, is not to be taken as an expression of opinion as to whether
or not they are subject to proprietary rights.
Printed in the United States of America.
9 8 7 6 5 4 3 2 1
SPIN 10697639
www.springer-ny.com
Springer-Verlag New York Berlin Heidelberg
A member of BertelsmannSpringer Science+Business Media GmbH
Arthur N. Popper
Departments of Psychology and
Department of Biology and
Neuroscience
Neuroscience and
Brown University
Cognitive Science Program
Box 1853
University of Maryland
Providence RI 02912, USA
College Park, MD 20742-4415, USA
Andrea_Simmons@brown.edu
AP17@umail.umd.edu
Richard R. Fay
Department of Psychology and
Parmly Hearing Institute
Loyola University of Chicago
Chicago, IL 60626, USA
rfay@wpo.it.luc.edu
Series Editors: Richard R. Fay and Arthur N. Popper
Cover illustration: Courtesy of Daniel Baleckaitis and Daniel Margoliash.
Library of Congress Cataloging-in-Publication Data
Acoustic communication / editors, Andrea Megela Simmons, Arthur N. Popper, Richard
R. Fay.
p. cm.—(Springer handbook of auditory research; 16)
Includes bibliographical references.
ISBN 0-387-98661-8 (alk. paper)
1. Animal sounds. 2. Sound production by animals. 3. Animal communication.
I. Simmons, Andrea Megela. II. Popper, Arthur N. III. Fay, Richard R. IV. Springer
handbook of auditory research; v. 16.
QL765 .A32 2002
591.59¢4—dc21
2001057680
ISBN 0-387-98661-8
Printed on acid-free paper.
© 2003 Springer-Verlag New York, Inc.
All rights reserved. This work may not be translated or copied in whole or in part without the
written permission of the publisher (Springer-Verlag New York, Inc., 175 Fifth Avenue, New
York, NY 10010, USA), except for brief excerpts in connection with reviews or scholarly
analysis. Use in connection with any form of information storage and retrieval, electronic
adaptation, computer software, or by similar or dissimilar methodology now know or hereafter developed is forbidden.
The use in this publication of trade names, trademarks, service marks, and similar terms, even
if they are not identified as such, is not to be taken as an expression of opinion as to whether
or not they are subject to proprietary rights.
Printed in the United States of America.
9 8 7 6 5 4 3 2 1
SPIN 10697639
www.springer-ny.com
Springer-Verlag New York Berlin Heidelberg
A member of BertelsmannSpringer Science+Business Media GmbH