Contents
1
The Eutrophication Problem in Temperate Lakes:
Practical Aspects and Theoretical Ramifications. . . . . . . . . . .. 1
1.1
Eutrophication: Consequences and Correctives. . . . . . . . . . . .. 2
1.2 Direct and Indirect Effects of Herbivorous Zooplankton. . . . . .. 5
1.3 Stability and Persistence in Grazer-Controlled Systems. . . . . . .. 8
1.4 Scope and Strategy . . . . • . . . . . • . . . . . . . . . . . . . . . • . . . • 10
2
The Biogeochemical Theatre -
Phosphorus Cycling and Phosphorus Household in Lakes
13
2.1
Phosphorus Mass-Balance Models ..................... 14
2.2 Phosphorus Retention in Lakes ....................... 15
2.3 Biologically Mediated Phosphorus Retention Processes . • . . . . . 18
2.4 Partitioning of Phosphorus in the Plankton Community .••.... 21
2.5 The Dynamics of the Phosphorus Cycle in Lakes. . . . . . . . . . . . 25
2.6 Summary and Conclusions . . . . . . . . . . . . . . . . . . . . . . . . . . 30
2.7 Symbols, Definitions, and Units .•..................... 32
3
Algae and Nutrients: Uptake and Utilization of Limiting
Nutrients in Generalized Phytoplankton Species. . . . . . . . . . . . 33
3.1
Growth and Nutrient Utilization ...................... 33
3.2 Nutrient Uptake. ...........................•.... 35
3.3 Balancing Nutrient Uptake and Growth ................•. 37
3.4 Balancing Photosynthesis and Growth. . . . . . . . . . . . . . . . . . . 39
3.5 Phytoplankton Growth Model Parametrization. . . . . . . • . . . . . 40
3.6 Resource Competition Under Phosphorus Limitation ........ 48
3.7 More Than One Limiting Nutrient - P and N Limitation ....... 51
3.8 Summary and Conclusions. . . . . . . . . . . . . . . . . . . . . . . . . . 59
3.9 Symbols, Definitions, and Units ....................... 60
4
Herbivores and Algae: Food Utilization, Growth
and Reproduction in Generalist Filter Feeders . . . . . . . . . . . . . 63
4.1
Characteristics of Daphnia Biology. . . . . . . . . . . . . . . . . . . . . 64