56
J. Can adell, A. Djema, B. L6pez, F. L., S. Sabate, D. Siscart and C.A. Gracia
600
600
500
500
C)I
400
400
to
E
0
~ 300
300
:E
0
3
0> 200
rI.
200
'<
100
100
0
0
Root
Root
Biomass
Production
15000
15000
Leaves
~
,
>. 10000
C)I
10000
A
E
0
ro
D>
u 5000
3
~
5000
' "
O -'----L--o
Maintenance cost
Formation cost
Fig. 4.3. Above: Fine root biomass and annual production; below: annual metabolic cost of fine
root formation and maintenance compared with formation and maintenance costs of leaves
References
Aber TO, Melillo JM, Nadelhoffer KJ, McClaugherty CA, Pastor J (1985) Fine root turnover in forest ecosystems in relation to quantity and form of nitrogen availability: a comparision of two
methods. Oecologia 66:317-321
Agren GI, Axelsson B, Flower-Ellis JGK, Linder S, Persson H, Staaf H, Troeng E (1980) Annual
carbon budget for a young Scots pine. Ecol Bull 32:307-313
Burke MK, Raynal DJ (1994) Fine root growth phenology, production, and turnover in a northern hardwood forest ecosystem. Plant Soil 162:135-146
Caldwell MM, Fernandez OA (1975) Dynamics of Great Basin shrub root systems. In: Hadley NF
(ed) Environmental physiology of desert organisms. Halstead Press, New York, pp 38-51
Canadell J, L6pez-Soria L (1998) Lignotuber reserves support regrowth following clipping of
two Mediterranean shrubs. Funct EcoI12:31-38