178
Carlos A. Gracia, Estibaliz Tello, Santiago Sabate and Juan Bellot
increases due to the rise in CO2 (Mooney et al. 1991; Skiles and Hanson 1994;
Medlyn and Dewar 1996). In holm oak forests water and temperature stresses
are limiting factors and their effects can overcome the fertilization effect of
CO2 increase.
Holm oak forests are highly sensitive to the increasing temperature due to
climate change and, even if WUE increases, most of the increased carbon
gain is accumulated in the soil due to higher litterfall and mortality (for
control and minimum thinning treatments). This situation would be less
critical for thinning treatments, because there is a higher water availability
and the respiration increase is proportionally lower than that in control
treatments. Thinning promotes a slight attenuation of the negative effects of
climate change, and could be a useful tool for improving the response to the
potential hydrological stress induced by climate change.
References
Albeza E, Sabate S, Escarre A, Gracia CA (1995) A long-term thinning experiment on a Quercus
ilex L. forest: changes in leaf characteristics and dynamics in response to different treatment
intensities. In: Jenkins A, Ferrier RC, Kirby C (eds) Ecosystem manipulation experiments.
Ecosystems Research Report 20, Commission of the European Communities, Luxembourg,
pp200-208
Bellot J (1989) AmUisis de los flujos de la deposici6n global, trascolaci6n, escorrentia cortical y
deposici6n Seca en el encinar mediternineo de L'Avic (Sierra de Prades, Tarragona). PhD
Thesis, University of Alicante, Alicante
Bellot J, Sanchez JR, Lled6 MJ, Martinez P, Escarre A (1992) Litterfall as a measure of primary
production in Mediterranean holm-oak forest. Vegetatio 99/100:69-76
Djema A, L6pez B, Sabate S, Gracia C (1994) A long-term thinning experiment on a Quercus ilex
L. forest: changes in root biomass and nutrient contents. Not Bioi (Santiago de Chile) 2(3):21
Kummerow J,Ellis BA (1989) Structure and function in chaparral shrubs. In: Keeley SC (ed) The
California chaparral. Paradigms reexamined. Natural History Museum of Los Angeles
County, Los Angeles,pp 141-150
Lled6 MJ, Sanchez JR, Bellot J, Boronat J, Ibanez H, Escarre A (1992) Structure biomass and production of a resprouted holm-oak (Quercus ilex) forest in NE Spain. Vegetatio 99/100:51-59
McKenney MS, Rosenberg NJ (1993) Sensitivity of some potential evapotranspiration estimation methods to climate change. Agric For Meteorol 64:81-110
Medlyn BE, Dewar RC (1996) A model of long-term response of carbon allocation and productivity of forests to increased CO2 concentration and nitrogen deposition. Global Change Bioi
2:367-376
Mooney HA, Drake BG, Luxmore RJ, Oechel WC, Pitelka LF (1991) Predicting ecosystem responses to elevated CO2 concentration. BioScience 41:96-104
Pinol J, Lled6 MJ, Escarre A (1991) Hydrological balance of two Mediterranean forested catchments (Prades, northeast Spain). Hydrol Sci J 36:95-107
Rotmans J, Hulme M, Downing IE (1994) Climate change implications for Europe. An application of the ESCAPE model. Global Environ Change 4:97-124
Sabate S (1993) Canopy structure and nutrient content in a Quercus ilex L. forest of Prades
mountains: effect of natural and experimental manipulation of growth conditions. PhD Thesis, University of Barcelona, Barcelona
Sala A (1992) Water relations, canopy structure, and canopy gas exchange in a Quercus ilex forest: variations in time and space. PhD Thesis, University of Barcelona, Barcelona
Carlos A. Gracia, Estibaliz Tello, Santiago Sabate and Juan Bellot
increases due to the rise in CO2 (Mooney et al. 1991; Skiles and Hanson 1994;
Medlyn and Dewar 1996). In holm oak forests water and temperature stresses
are limiting factors and their effects can overcome the fertilization effect of
CO2 increase.
Holm oak forests are highly sensitive to the increasing temperature due to
climate change and, even if WUE increases, most of the increased carbon
gain is accumulated in the soil due to higher litterfall and mortality (for
control and minimum thinning treatments). This situation would be less
critical for thinning treatments, because there is a higher water availability
and the respiration increase is proportionally lower than that in control
treatments. Thinning promotes a slight attenuation of the negative effects of
climate change, and could be a useful tool for improving the response to the
potential hydrological stress induced by climate change.
References
Albeza E, Sabate S, Escarre A, Gracia CA (1995) A long-term thinning experiment on a Quercus
ilex L. forest: changes in leaf characteristics and dynamics in response to different treatment
intensities. In: Jenkins A, Ferrier RC, Kirby C (eds) Ecosystem manipulation experiments.
Ecosystems Research Report 20, Commission of the European Communities, Luxembourg,
pp200-208
Bellot J (1989) AmUisis de los flujos de la deposici6n global, trascolaci6n, escorrentia cortical y
deposici6n Seca en el encinar mediternineo de L'Avic (Sierra de Prades, Tarragona). PhD
Thesis, University of Alicante, Alicante
Bellot J, Sanchez JR, Lled6 MJ, Martinez P, Escarre A (1992) Litterfall as a measure of primary
production in Mediterranean holm-oak forest. Vegetatio 99/100:69-76
Djema A, L6pez B, Sabate S, Gracia C (1994) A long-term thinning experiment on a Quercus ilex
L. forest: changes in root biomass and nutrient contents. Not Bioi (Santiago de Chile) 2(3):21
Kummerow J,Ellis BA (1989) Structure and function in chaparral shrubs. In: Keeley SC (ed) The
California chaparral. Paradigms reexamined. Natural History Museum of Los Angeles
County, Los Angeles,pp 141-150
Lled6 MJ, Sanchez JR, Bellot J, Boronat J, Ibanez H, Escarre A (1992) Structure biomass and production of a resprouted holm-oak (Quercus ilex) forest in NE Spain. Vegetatio 99/100:51-59
McKenney MS, Rosenberg NJ (1993) Sensitivity of some potential evapotranspiration estimation methods to climate change. Agric For Meteorol 64:81-110
Medlyn BE, Dewar RC (1996) A model of long-term response of carbon allocation and productivity of forests to increased CO2 concentration and nitrogen deposition. Global Change Bioi
2:367-376
Mooney HA, Drake BG, Luxmore RJ, Oechel WC, Pitelka LF (1991) Predicting ecosystem responses to elevated CO2 concentration. BioScience 41:96-104
Pinol J, Lled6 MJ, Escarre A (1991) Hydrological balance of two Mediterranean forested catchments (Prades, northeast Spain). Hydrol Sci J 36:95-107
Rotmans J, Hulme M, Downing IE (1994) Climate change implications for Europe. An application of the ESCAPE model. Global Environ Change 4:97-124
Sabate S (1993) Canopy structure and nutrient content in a Quercus ilex L. forest of Prades
mountains: effect of natural and experimental manipulation of growth conditions. PhD Thesis, University of Barcelona, Barcelona
Sala A (1992) Water relations, canopy structure, and canopy gas exchange in a Quercus ilex forest: variations in time and space. PhD Thesis, University of Barcelona, Barcelona