O’Farrell, P. J., & Milton, S. J. (2006). Road verge and rangeland plant communities in the
southern Karoo: Exploring what influences diversity, dominance and cover. Biodiversity and
Conservation, 15, 921–938.
Pellissier, V., Roze, F., Aguejdad, R., Quenol, H., & Clergeau, P. (2008). Relationships between
soil seed bank, vegetation and soil fertility along an urbanisation gradient. Applied Vegetation
Science, 11, 325–334.
Penone, C., Kerbiriou, C., Julien, J.-F., Julliard, R., Machon, N., & Le Viol, I. (2013).
Urbanisation effect on Orthoptera: Which scale matters? Insect Conservation and Diversity, 6,
319–327.
Penone, C., Machon, N., Julliard, R., & Le Viol, I. (2012). Do railway edges provide functional
connectivity for plant communities in an urban context? Biological Conservation, 148,
126–133.
Ricketts, T. H. (2001). The matrix matters: Effective isolation in fragmented landscapes. The
American Naturalist, 158, 87–99.
Ridding, L. E., Redhead, J. W., & Pywell, R. F. (2015). Fate of semi-natural grassland in England
between 1960 and 2013: A test of national conservation policy. Global Ecology and
Conservation, 4, 516–525.
Ries, L., Debinski, D. M., & Wieland, M. L. (2001). Conservation value of roadside prairie
restoration to butterfly communities. Conservation Biology, 15, 401–411.
Roy, V., & de Blois, S. (2006). Using functional traits to assess the role of hedgerow corridors as
environmental filters for forest herbs. Biological Conservation, 130, 592–603.
Russ, J. M., & Montgomery, W. I. (2002). Habitat associations of bats in Northern Ireland:
Implications for conservation. Biological Conservation, 108, 49–58.
Russo, D., & Jones, G. (2003). Use of foraging habitats by bats in a Mediterranean area determined
by acoustic surveys: Conservation implications. Ecography, 26, 197–209.
Saarinen, K., Valtonen, A., Jantunen, J., & Saarnio, S. (2005). Butterflies and diurnal moths along
road verges: Does road type affect diversity and abundance? Biological Conservation, 123,
403–412.
Stenhouse, R. N. (2004). Fragmentation and internal disturbance of native vegetation reserves in
the Perth metropolitan area, Western Australia. Landscape and Urban Planning, 68, 389–401.
Taylor, P. D., Fahrig, L., Henein, K., & Merriam, G. (1993). Connectivity is a vital element of
landscape structure. Oikos, 68, 571–573.
Temple, H. J., & Terry, A. (2009). The status and distribution of european mammals.
Luxembourg: Office for Official Publications of the European Communities.
Theuerkauf, J., & Rouys, S. (2006). Do Orthoptera need human land use in Central Europe? The
role of habitat patch size and linear corridors in the Biaowieza forest, Poland. Biodiversity and
Conservation, 15, 1497–1508.
Tikka, P. M., Hogmander, H., & Koski, P. S. (2001). Road and railway verges serve as dispersal
corridors for grassland plants. Landscape Ecology, 16, 659–666.
Vandevelde, J.-C., Bouhours, A., Julien, J.-F., Couvet, D., & Kerbiriou, C. (2014). Activity of
European common bats along railway verges. Ecological Engineering, 64, 49–56.
Vandevelde, J.-C., Penone, C., & Julliard, R. (2012). High-speed railways are not barriers to
Pyronia tithonus butterfly movements. Journal of Insect Conservation, 16, 801–803.
Verboom, B., & Huitema, H. (1997). The importance of linear landscape elements for the
pipistrelle Pipistrellus pipistrellus and the serotine bat Eptesicus serotinus. Landscape Ecology,
12, 117–125.
Verboom, B., & Spoelstra, K. (1999). Effects of food abundance and wind on the use of tree
lines by an insectivorous bat, Pipistrellus pipistrellus. Canadian Journal of Zoology, 77, 1393–
1401.
Vergnes, A., Le Viol, I., & Clergeau, P. (2012). Green corridors in urban landscapes affect the
arthropod communities of domestic gardens. Biological Conservation, 145, 171–178.
Voisin, J. F. (2003). Atlas des Orthoptères et des Mantides de France [Atlas of French Orthoptera
and Mantis of France]. Paris: MNHN, Collection Patrimoines Naturels.
16 Ecological Roles of Railway Verges in Anthropogenic Landscapes …
275
southern Karoo: Exploring what influences diversity, dominance and cover. Biodiversity and
Conservation, 15, 921–938.
Pellissier, V., Roze, F., Aguejdad, R., Quenol, H., & Clergeau, P. (2008). Relationships between
soil seed bank, vegetation and soil fertility along an urbanisation gradient. Applied Vegetation
Science, 11, 325–334.
Penone, C., Kerbiriou, C., Julien, J.-F., Julliard, R., Machon, N., & Le Viol, I. (2013).
Urbanisation effect on Orthoptera: Which scale matters? Insect Conservation and Diversity, 6,
319–327.
Penone, C., Machon, N., Julliard, R., & Le Viol, I. (2012). Do railway edges provide functional
connectivity for plant communities in an urban context? Biological Conservation, 148,
126–133.
Ricketts, T. H. (2001). The matrix matters: Effective isolation in fragmented landscapes. The
American Naturalist, 158, 87–99.
Ridding, L. E., Redhead, J. W., & Pywell, R. F. (2015). Fate of semi-natural grassland in England
between 1960 and 2013: A test of national conservation policy. Global Ecology and
Conservation, 4, 516–525.
Ries, L., Debinski, D. M., & Wieland, M. L. (2001). Conservation value of roadside prairie
restoration to butterfly communities. Conservation Biology, 15, 401–411.
Roy, V., & de Blois, S. (2006). Using functional traits to assess the role of hedgerow corridors as
environmental filters for forest herbs. Biological Conservation, 130, 592–603.
Russ, J. M., & Montgomery, W. I. (2002). Habitat associations of bats in Northern Ireland:
Implications for conservation. Biological Conservation, 108, 49–58.
Russo, D., & Jones, G. (2003). Use of foraging habitats by bats in a Mediterranean area determined
by acoustic surveys: Conservation implications. Ecography, 26, 197–209.
Saarinen, K., Valtonen, A., Jantunen, J., & Saarnio, S. (2005). Butterflies and diurnal moths along
road verges: Does road type affect diversity and abundance? Biological Conservation, 123,
403–412.
Stenhouse, R. N. (2004). Fragmentation and internal disturbance of native vegetation reserves in
the Perth metropolitan area, Western Australia. Landscape and Urban Planning, 68, 389–401.
Taylor, P. D., Fahrig, L., Henein, K., & Merriam, G. (1993). Connectivity is a vital element of
landscape structure. Oikos, 68, 571–573.
Temple, H. J., & Terry, A. (2009). The status and distribution of european mammals.
Luxembourg: Office for Official Publications of the European Communities.
Theuerkauf, J., & Rouys, S. (2006). Do Orthoptera need human land use in Central Europe? The
role of habitat patch size and linear corridors in the Biaowieza forest, Poland. Biodiversity and
Conservation, 15, 1497–1508.
Tikka, P. M., Hogmander, H., & Koski, P. S. (2001). Road and railway verges serve as dispersal
corridors for grassland plants. Landscape Ecology, 16, 659–666.
Vandevelde, J.-C., Bouhours, A., Julien, J.-F., Couvet, D., & Kerbiriou, C. (2014). Activity of
European common bats along railway verges. Ecological Engineering, 64, 49–56.
Vandevelde, J.-C., Penone, C., & Julliard, R. (2012). High-speed railways are not barriers to
Pyronia tithonus butterfly movements. Journal of Insect Conservation, 16, 801–803.
Verboom, B., & Huitema, H. (1997). The importance of linear landscape elements for the
pipistrelle Pipistrellus pipistrellus and the serotine bat Eptesicus serotinus. Landscape Ecology,
12, 117–125.
Verboom, B., & Spoelstra, K. (1999). Effects of food abundance and wind on the use of tree
lines by an insectivorous bat, Pipistrellus pipistrellus. Canadian Journal of Zoology, 77, 1393–
1401.
Vergnes, A., Le Viol, I., & Clergeau, P. (2012). Green corridors in urban landscapes affect the
arthropod communities of domestic gardens. Biological Conservation, 145, 171–178.
Voisin, J. F. (2003). Atlas des Orthoptères et des Mantides de France [Atlas of French Orthoptera
and Mantis of France]. Paris: MNHN, Collection Patrimoines Naturels.
16 Ecological Roles of Railway Verges in Anthropogenic Landscapes …
275