343
Kerr PH, Winterton SL (2008) Do parasitic flies attack mites? Evidence in Baltic amber. Biol J
Linn Soc 93:9–13
Khaustov AA, Perkovsky EE (2010) The first fossil record of mites of the family Pyemotidae
(Acari: Heterostigmata), with description of a new species from Rovno Amber. Palaeontol J
44:418–421
Khaustov AA, Poinar GO Jr (2010) Protoresinacarus brevipedis gen. n., sp. n. from Early
Cretaceous Burmese amber: the first fossil record of mites of the family Resinacaridae (Acari:
Heterostigmata: Pyemotoidea). Hist Biol 23:219–222
Klimov PB, OConnor BM, Chetverikov PE, Bolton SJ, Pepato AR, Mortazavi AL, Tolstikov AV,
Bauchan GR, Ochoa R (2018) Comprehensive phylogeny of acariform mites (Acariformes)
provides insights on the origin of the four-legged mites (Eriophyoidea), a long branch. Mol
Phylogenet Evol 119:105–117
Klompen H (2010) Holothyrids and ticks: new insights from larval morphology and DNA sequencing, with the description of a new species of Diplothyrus (Parasitiformes: Neothyridae).
Acarologia 50:269–285
Klompen H, Grimaldi D (2001) First Mesozoic record of a parasitiform mite: a larval argasid tick
in Cretaceous amber (Acari: Ixodida: Argasidae). Ann Entomol Soc Am 94:10–15
Klompen JSH, Black WC, Keirans JE, Oliver JH Jr (1996) Evolution of ticks. Annu Rev Entomol
41:141–161
Klompmaker AA, Chistoserdov AY, Felder DL (2016) Possible shell disease in 100 million-yearold crabs. Dis Aquat Org 119:91–99
Konishi K, Shimazaki K (1998) Halarachnid mites infesting the respiratory tract of Stellar sea
lions. Biosphere Cons 1:45–48
Krantz GW (2009) Habits and habitats. In: Krantz GW, Walter DE (eds) A manual of acarology,
3rd edn. Texas Tech University Press, Lubbock, pp 64–82
Labandeira CC, Li L (2021). The history of insect parasitism and the mid-Mesozoic parasitoid
revolution. In: De Baets K, Huntley JW (eds) The Evolution and Fossil Record Of Parasitism:
Identification and Macroevolution of Parasites. Topics in Geobiology 49
Lamsdell JC (2013) Revised systematics of Palaeozoic ‘horseshoe crabs’ and the myth of monophyletic Xiphosura. Zool J Linnean Soc 167:1–27
Lamsdell JC, Braddy SJ (2010) Cope’s Rule and Romer’s theory: patterns of diversity and gigantism in eurypterids and Palaeozoic vertebrates. Biol Lett 6:265–269
Lane RS, Poinar GO Jr (1986) First fossil tick (Acari: Ixodidae) in New World amber. Int J Acarol
12:75–78
Le Conte Y, Ellis M, Ritter W (2010) Varroa mites and honey bee health: can Varroa explain part
of the colony losses. Apidologie 41:353–363
Legg DA (2014) Sanctacaris uncata: the oldest chelicerate (Arthropoda). Naturwissenschaften
101:1065–1073
Leibovitz L, Lwebart GA (2003) Diseases and symbionts: vulnerability despite tough shells.
In: Shuster CN Jr, Barlow HB, Brockmann HJ (eds) The American horseshoe crab. Harvard
University Press, Cambridge, MA, pp 245–267
Leung TLF (2017) Fossils of parasites: what can the fossil record tell us about the evolution of
parasitism? Biol Rev 92:410–430
Lindquist EE (1986) The world genera of Tarsonemidae (Acari: Heterostigmata): a morphological, phylogenetic, and systematic revision, with a reclassification of family-group taxa in the
Heterostigmata. Mem Entomol Soc Canada 136:1–517
Lindquist EE, Krantz GW, Walter DE (2009) Classification. In: Krantz GW, Walter DE (eds) A
manual of acarology, 3rd edn. Texas Tech University Press, Lubbock, pp 97–103
Lozano-Fernandez J, Tanner AR, Giacomelli M, Carton R, Vinther J, Edgecombe GD, Pisani D
(2019) Increasing species sampling in chelicerate genomic-scale datasets provides support for
monophyly of Acari and Arachnida. Nat Commun 10:2295
Lyubarsky GY, Perkovsky EE (2012) The first Eocene species of the genus Cryptophagus
(Coleoptera, Clavicornia, Cryptophagidae). Vest Zool 46:36–40
Magowski WŁ (1994) Discovery of the first representative of the mite subcohort Heterostigmata
(Arachinida: Acari) in the Mesozoic Siberian amber. Acarologia 35:229–241
9 Chelicerates as Parasites
Kerr PH, Winterton SL (2008) Do parasitic flies attack mites? Evidence in Baltic amber. Biol J
Linn Soc 93:9–13
Khaustov AA, Perkovsky EE (2010) The first fossil record of mites of the family Pyemotidae
(Acari: Heterostigmata), with description of a new species from Rovno Amber. Palaeontol J
44:418–421
Khaustov AA, Poinar GO Jr (2010) Protoresinacarus brevipedis gen. n., sp. n. from Early
Cretaceous Burmese amber: the first fossil record of mites of the family Resinacaridae (Acari:
Heterostigmata: Pyemotoidea). Hist Biol 23:219–222
Klimov PB, OConnor BM, Chetverikov PE, Bolton SJ, Pepato AR, Mortazavi AL, Tolstikov AV,
Bauchan GR, Ochoa R (2018) Comprehensive phylogeny of acariform mites (Acariformes)
provides insights on the origin of the four-legged mites (Eriophyoidea), a long branch. Mol
Phylogenet Evol 119:105–117
Klompen H (2010) Holothyrids and ticks: new insights from larval morphology and DNA sequencing, with the description of a new species of Diplothyrus (Parasitiformes: Neothyridae).
Acarologia 50:269–285
Klompen H, Grimaldi D (2001) First Mesozoic record of a parasitiform mite: a larval argasid tick
in Cretaceous amber (Acari: Ixodida: Argasidae). Ann Entomol Soc Am 94:10–15
Klompen JSH, Black WC, Keirans JE, Oliver JH Jr (1996) Evolution of ticks. Annu Rev Entomol
41:141–161
Klompmaker AA, Chistoserdov AY, Felder DL (2016) Possible shell disease in 100 million-yearold crabs. Dis Aquat Org 119:91–99
Konishi K, Shimazaki K (1998) Halarachnid mites infesting the respiratory tract of Stellar sea
lions. Biosphere Cons 1:45–48
Krantz GW (2009) Habits and habitats. In: Krantz GW, Walter DE (eds) A manual of acarology,
3rd edn. Texas Tech University Press, Lubbock, pp 64–82
Labandeira CC, Li L (2021). The history of insect parasitism and the mid-Mesozoic parasitoid
revolution. In: De Baets K, Huntley JW (eds) The Evolution and Fossil Record Of Parasitism:
Identification and Macroevolution of Parasites. Topics in Geobiology 49
Lamsdell JC (2013) Revised systematics of Palaeozoic ‘horseshoe crabs’ and the myth of monophyletic Xiphosura. Zool J Linnean Soc 167:1–27
Lamsdell JC, Braddy SJ (2010) Cope’s Rule and Romer’s theory: patterns of diversity and gigantism in eurypterids and Palaeozoic vertebrates. Biol Lett 6:265–269
Lane RS, Poinar GO Jr (1986) First fossil tick (Acari: Ixodidae) in New World amber. Int J Acarol
12:75–78
Le Conte Y, Ellis M, Ritter W (2010) Varroa mites and honey bee health: can Varroa explain part
of the colony losses. Apidologie 41:353–363
Legg DA (2014) Sanctacaris uncata: the oldest chelicerate (Arthropoda). Naturwissenschaften
101:1065–1073
Leibovitz L, Lwebart GA (2003) Diseases and symbionts: vulnerability despite tough shells.
In: Shuster CN Jr, Barlow HB, Brockmann HJ (eds) The American horseshoe crab. Harvard
University Press, Cambridge, MA, pp 245–267
Leung TLF (2017) Fossils of parasites: what can the fossil record tell us about the evolution of
parasitism? Biol Rev 92:410–430
Lindquist EE (1986) The world genera of Tarsonemidae (Acari: Heterostigmata): a morphological, phylogenetic, and systematic revision, with a reclassification of family-group taxa in the
Heterostigmata. Mem Entomol Soc Canada 136:1–517
Lindquist EE, Krantz GW, Walter DE (2009) Classification. In: Krantz GW, Walter DE (eds) A
manual of acarology, 3rd edn. Texas Tech University Press, Lubbock, pp 97–103
Lozano-Fernandez J, Tanner AR, Giacomelli M, Carton R, Vinther J, Edgecombe GD, Pisani D
(2019) Increasing species sampling in chelicerate genomic-scale datasets provides support for
monophyly of Acari and Arachnida. Nat Commun 10:2295
Lyubarsky GY, Perkovsky EE (2012) The first Eocene species of the genus Cryptophagus
(Coleoptera, Clavicornia, Cryptophagidae). Vest Zool 46:36–40
Magowski WŁ (1994) Discovery of the first representative of the mite subcohort Heterostigmata
(Arachinida: Acari) in the Mesozoic Siberian amber. Acarologia 35:229–241
9 Chelicerates as Parasites